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Left 4 Dead puts four human playable or AI controlled survivors of an apocalyptic pandemic against hordes of aggressive "Infected" zombies in a cinematic backdrop in which the players "star" as each character in a fictional movie. The Survivors goal is to help each other make it through various scenarios to safety. The Infected are controlled by an AI that dynamically balances difficulty and mood depending on the players progress and situation. In an alternate game mode, human players can control up to four different monsters with special abilities and cooperate to stop the Survivor players.

It was released on November 18, 2008 in the United States and November 21, 2008 in Europe to coincide with the tenth anniversary of the release of Half-Life. A five-minute trailer was released on Halloween. A public demo was released on November 6 for pre-purchasers and was released for the general public on November 11 for both systems. The demo contains the first two out of five maps of the "No Mercy" campaign and is playable in both single player and multi-player. Left 4 Dead went gold on November 13, 2008.